Dna single molecules adsorbed on mica: conformational fluctuations and recognition processes studied by Scanning Force Microscope

Bruno Samori', Giampaolo Zuccheri
 
 Dipartimento di Biochimica. Universita' di Bologna
via Irnerio 48, 40126 Bologna (Italy)
 

Chemists used to sample Avogadro numbers of molecules for their experiments. The properties they measured were averaged on the entire ensemble of those molecules and over all conformations accessible to them. A new exciting dimension has been opened: a single molecule can be simultaneously touched and observed, and then also dissected and mechanically manipulated to drive its structure towards new conformations and states that otherwise can be very hard to reach. Conformational rearrangements from these states and thermal fluctuations of supercoiled and linear DNA molecules were followed and observed in fluid, in real time, with a Scanning Force Microscope (SFM). Local curvatures and flexibilities of the chains were monitored and analyzed at the single molecule level and their modulation by the sequence was studied. The spatial resolution of SFM settles this analysis in the nanometer scale. This is the length scale of the long-range recognition of conformational properties and dynamics labeled "indirect-read out". This recognition mechanism of not-so-local conformational properties is used by the DNA-tracking-proteins during their search of specific binding sites along the DNA chain. This work shows how topics of polymer chemistry and biology can be transferred to the single-molecule level.
 

The capability of controlling the adsorption and reactivity of single dna molecules on surfaces opens the perspective of creating a novel class of nanotechnologies of the nucleic acids.

 

References

 G. Zuccheri, R. Thei Dame, M. Aquila, I. Muzzalupo, B. Samori', Conformational fluctuations of supercoiled DNA molecules observed in real time with a scanning force microscope, Applied Physics A 1998, 66, S585-S589

B. Samori',  Stretching, tearing and dissecting single molecules of DNA, Angew. Chem. Int. Ed. Engl., 1998, 37, 2198-2200